Gramma, the beloved Galapagos tortoise and the oldest resident of the San Diego Zoo, has passed away at the estimated age of 141. Known for her gentle demeanor, Gramma lived through significant historical events, including two world wars and the presidencies of over 20 U.S. leaders. She was not only a remarkable testament to longevity but also a cherished figure among visitors for almost a century. The zoo is mourning the loss of its longest-lived resident, who outlived empires and became a symbol of resilience and joy for many.